Here is a list of my publications. Please follow the links to obtain the pdf files.

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13) Sherley, R. B., Barham, B. J., Barham, P. J., Campbell, K. J., Crawford, R. J. M., Grigg, J., Horswill, C., McInnes, A. MorrisT. L., Pichegru, L., Steinfurth, A., Weller, F., Winker, H., Votier, S. C. (2018). Bayesian inference reveals positive but subtle effects of experimental fishery closures on marine predator demographics. Proceedings of the Royal Society B: Biological Sciences, 285(1871).


12) Horswill C., Trathan. P. N., Ratcliffe, N. 2017. Linking extreme interannual changes in prey availability to foraging behaviour and breeding investment in a marine predator, the macaroni penguin. PLoS ONE 12(9): e0184114.

11) Thaxter, C.B., Horswill, C., Ross, K.E., Austin, G.E., Balmer, D.E. Niall, H.K. 2017. Urban breeding gull surveys: A survey design simulation. BTO Research Report No. 699. British Trust for Ornithology, Thetford.

10) Horswill, C., O’Brien, S. H., Robinson R. A. 2017. Density-dependence and marine bird populations: are wind farm assessments precautionary? Journal of Applied Ecology. 54, 1406–1414 doi: 10.1111/1365-2664.12841


9) Horswill, C., Walker R. H., Humphreys E. M., Robinson R. A. 2016. Review of mark-recapture studies on UK seabirds that are run through the BTO’s Retrapping Adults for Survival (RAS) network. JNCC Report No. 600. Joint Nature Conservation Committee, Peterborough.

8) Horswill, C., Ratcliffe, N., Green, J.A., Phillips, R.A., Trathan, P.N., Matthiopoulos, J., 2016. Unravelling the relative roles of top-down and bottom-up forces driving population change in an oceanic predator. Ecology 97: 1919-1928. doi:10.1002/ecy.1452

7) Horswill, C., Matthiopoulos, J., Ratcliffe, N., Green, J.A., Trathan, P.N., McGill, R.A.R., Phillips, R.A., O’Connell, T.C., 2016. Drivers of intrapopulation variation in resource use in a generalist predator, the macaroni penguin. Marine Ecology Progress Series 548, 223–247. doi:10.3354/meps11626


6) Horswill, C. and Robrinson, R. 2015. Review of seabird demographic rates and density dependence. JNCC Report No. 552. Joint Nature Conservation Committee, Peterborough.

5) Horswill, C. The relative importance of opposing drivers in determining population change in macaroni penguins Eudyptes chrysolophus. PhD Thesis. British Antarctic Survey, University of Liverpool and University of Glasgow.

4) Ratcliffe, N., Hill, S.L., Staniland, I.J., Brown, R., Adlard, S., Horswill, C., Trathan, P.N., 2015. Do krill fisheries compete with macaroni penguins? Spatial overlap in prey consumption and catches during winter. Diversity and Distributions. doi:10.1111/ddi.12366


3) Ratcliffe, N., Crofts, S., Brown, R.M., Baylis, A.M.M., Adlard, S., Horswill, C., Venables, H., Taylor, P., Trathan, P.N., Staniland, I.J., 2014. Love thy neighbour or opposites attract? Patterns of spatial segregation and association among crested penguin populations during winter. Journal of Biogeography 41, 1183–1192. doi:10.1111/jbi.12279

2)  Horswill, C., Matthiopoulos, J., Green, J.A., Meredith, M.P., Forcada, J., Peat, H.J., Preston, M., Trathan, P.N., Ratcliffe, N., 2014. Survival in macaroni penguins and the relative importance of different drivers: individual traits, predation pressure and environmental variability. Journal of Animal Ecology 83, 1057–1067. doi:10.1111/1365-2656.12229


1) Horswill, C., Jackson, J.A., 2012. Humpback whales wintering at Pitcairn Island, South Pacific. Marine Biodiversity Records 5. doi:10.1017/S1755267212000693